What is the question "Mah Ne'enchah Behemah?
Malbim: The Navi asks, if man sinned what sin did animals do? Why are animals groaning, and lack pasture?
What Behemah is discussed?
Radak: This is the Klal; afterwards it details (cattle and flock).
What is the meaning of "Navochu"?
Rashi: They were entangled and erred in forests and Midbaros.
Radak: This is confusion, like one who is confused, and does not know what to do. This is like "Nevuchim Hem ba'Aretz" (Shemos 14:3).
Malbim: Animals are confused because they have no pasture.
What is the significance of "Gam Edrei Tzon"?
Radak: Sometimes flocks find pasture that cattle do not find, because flocks ascend mountains and hills; cattle usually do not.
Malbim: Did also the flocks sin, and this is why they are punished?!
What do we learn from "Neshamu"?
Rashi: They were ruined in their Ashmah (guilt).
Radak: This is desolation. The same applies to "Tesham Shomron" (Hoshe'a 14:1).
Malbim: Refer to 1:18:4:4.