What is "Kodesh" in this Pasuk referring to?
Rashi and Rashbam: Here too, it is referring to Terumah.
What exactly, is the Zar who ate Kodesh be'Shogeg obligated to pay the Kohen (See Sifsei Chachamim)?
Rashi and Ramban: He is obligated to pay (not with money, but) with Chulin-fruit that adopts Kedushas Terumah. 1
What happens to a Zar who eats Terumah be'Meizid?
Rashi (Shabbos, 69b): He is not Chayav to add a fifth but is Chayav Misah bi'Yedei Shamayim 1 and does not add
Abaye (Kesuvos 30a) holds that R. Nechunya ben Hakanah exempts from payment for Aveiros of Misah bi'Ydei Shamayim from payment, and even if he was Shogeg. If so, why does a Zar pay for eating Terumah?
Riva citing R"M of Kutzi: The exemption does not apply to matters of Kaparah.